Morphological analysis of the spores and their bearing on the taxonomy and evolution of Pteridaceae have been dealt with. The spore morphological combinations in evolutionary successions are trilete-non-perinous, trilete-perinous, monolete-non-perinous and monolete-perinous. It is suggested that the levels of evolutionary progress of the various taxa are in the order of succession, Pteroids, Gymnogrammoids and Lindsaeoids, but each have evolved along parallel lines from the Dicksonioid-Dennstaedtioid complex.