The Lachlan Fold Belt has the velocity‐depth structure of continental crust, with a thickness exceeding 50 km under the region of highest topography in Australia, and in the range 41–44 km under the central Fold Belt and Sydney Basin. There is no evidence of high upper crustal velocities normally associated with marginal or back‐arc basin crustal rocks. The velocities in the lower crust are consistent with an overall increase in metamorphic grade and/or mafic mineral content with depth. Continuing tectonic development throughout the region and the negligible seismicity at depths greater than 30 km indicate that the lower crust is undergoing ductile deformation.