SUMMARYThe somatic chromosome complement inSynedrella nodiflorawas found to be 40, of which three pairs are satellited. The karyotypes have been analysed. The presence of 20 chromosomes was noted in diakinesis and metaphase I. Secondary association has been noted. The tapetal cells are quadrinucleate and a periplasmodium is formed.The pollen grains are trinucleate, echinate, ellipsoidal and tetracolporate. The sperm cells are elongated and rod‐shaped.The development of the female gametophyte is of the ‘Polygonum type’. The anti‐. podals are generally two in number, the upper cell being binucleate. In rare cases three uninucleate antipodal cells have also been observed. Endosperm formation is of the nuclear type. The embryo development conforms to theAsteradtype and agrees more or less with theSenecio variationof Johansen.The fruits derived from the disk florets are tubular and have two to three long pointed processes, while those from the ray florets are broad with numerous hairy outgrowths. Both types are