AbstractWe used 0.5 m diameter nets to sample the drift downstream from a peaking hydroelectric project on the Caney Fork River, Tennessee. The relative contribution of the terrestrial, lentic, and lotic environments to stream drift varied seasonally. Stream benthos (primarily Chironomidae andLirceus fontinalis) occurred in the drift throughout the year; however, their relative importance varied seasonally.Daphniasp. from the reservoir also occurred in the drift year round with the highest densities in fall and spring when they were the predominant species in the drift. In summer, terrestrial insects apparently utilized exposed portions of the river bed during periods of no power generation. In June as the river stage increased following initiation of generation, adult terrestrial insects became a major component of the drift. Threadfin shad (Dorosoma petenense) dominated the biomass in the drift during December. Although the importance of stream benthos to fish populations is generally recognized, the contribution of the terrestrial and upstream reservoir environments to the forage base in the tailwater may also be important and should be considered in developing management strategies for tailwater fisheries.