Coiled initials embedded in a stroma develop into multicellular, multinucleate ascogonia. Hyphae grow up around the ascogonia to form a prosenchymatous perithecial wall. As the ascocarp develops, an apical meristem produces cells that elongate downward into the centrum, forming long filaments, the apical paraphyses. The ascogonium produces multinucleate ascogenous cells which form a basal layer in the centrum. The ascogenous cells produce asci that grow up among the apical paraphyses, which disintegrate as the ascocarp matures. The asci are unitunicate, with a simple thickened cap punctuated by a pore. This pattern of development is typical of theNectria‐type.