SUMMARYSIBLEY, C.G., R. SCHODDE, and J.E. AHLQUIST. 1984. The relationships of the Australo-Papuan treecreepers Climacteridae as indicated by DNA-DNA hybridization. Emu 84: 236–241.The technique of DNA-DNA hybridization was used to compare the nuclear DNAs of the Brown TreecreeperClimacteris picumnusand the Rufous TreecreeperClimacteris rufawith the DNAs of other passerine taxa. The treecreepers were found to be the descendants of the oldest dichotomy (ca. 45–48 MYA) in the superfamily Menuroidea, a group that also includes the lyrebirds, scrub-birds, and bowerbirds. We recognize the family Climacteridae, superfamily Menuroidea, parvorder Corvi, suborder Passeres. Within the Climacteridae we recognize two genera,Climacterisfor the speciespicumnus, rufa, melanura, erythrops, andaffinis, andCormobatesforleucophaea(includingminor) andplacens.