Abstract1Although micro‐organisms are regarded as the principal agents of mineral cycling in soils, the role of bacteria has generally been emphasized, while that of fungi has been neglected.2Fungi are able to transform the majority of elementsin vitrobut whether they play an important role in soil is as yet unknown.3There is sufficient circumstantial evidence from soil studies to suggest that fungi may under certain conditions nitrify and oxidize reduced forms of sulphur. Their role in denitrification and sulphate reduction process is more speculative. While they appear incapable of nitrogen fixation, fungi undoubtedly play a major role in the mineralization of organic N,P, and S in soils, in the solubilization of insoluble phosphates, and participate in oxidation of manganese. Perhaps one of their most important roles is in the dissolution of silica and rocks thus releasing ions into the soil solution during weathering.4The ability of fungi to oxidize elementsin vitrodoes not compare with that of the chemoautotrophic bacteria. On the other handin vitroactivity tells us little about the activity of an organism in the soil.5At present our appreciation of the part played by fungi in mineral cycling in soils is limited by the techniques available, but there is little doubt that they have a major role to play in the cycling of elements other than carbo