Three seismically defined lower crustal blocks (LCBs) have been recognized to underlie the familiar tectonic‐stratigraphic zones of the Canadian Appalachians. Tectonic development of the orogen is inferred to have been strongly influenced by collision of the two outboard LCBs (Central and Avalon LCBs) against the sharply irregular North American craton (Grenville LCB), as evidenced by an abrupt offset in the trend of the orogen localized at this irregular edge. The LCBs are restored to a precollisional configuration by (1) removing Carboniferous strike‐slip motion on major faults linking through the Magdalen pull‐apart basin; and (2) removing Devonian(?) offset along a hypothesized fault through the present‐day Strait of Canso (Canso fault) to realign the Central and Avalon LCBs, presumed to have been continuous prior to accretion to North America. The restoration leads to (1) a qualitative explanation for deep seismic reflection observations across the Magdalen basin which suggest that portions of the underlying crust do not correspond to any of the three principal LCBs; and (2) a plate tectonic model for the orogen which suggests the backarc basin separating the Taconian (Ordovician) arc from the eastern margin of the early Paleozoic Iapetus ocean was at least as wide (>450 km) as the offset in the irregular Grenville LCB, inherited from rifting of Iapetus. The present‐day Southwest Newfoundland Transform Margin may be a segment of the Canso fault reactivated during Mesozoic opening of the Atlan