The megagametophyte ofPlumbagella micrantha, consisting of three cells: an egg, a central cell, and a single antipodal, represents the most structurally reduced female gametophyte of the flowering plants. In the absence of synergids, the pollen tube enters the megagametophyte through a filiform apparatus at the base of the egg, continues growth for 40μm through the egg‐central cell wall, until reaching a region of strong curvature near the chalazal end of the egg. A terminal pollen tube aperture then forms, the sperm cells and vegetative nucleus are released between the egg and central cell, and the sperm undergo gametic fusion, thereby effecting double fertilization. The mode of fertilization inPlumbagellais therefore similar to that ofPlumbagothat has been described elsewhere.