AbstractSeparate groups of non‐lactating cows and wether sheep grazed at similar herbage allowances for two successive 5‐d periods on swards that had previously been grazed frequently or infrequently with the intention of creating differences in canopy structure. Measurements were made of sward structure and composition, ingestive behaviour and diet composition. The preliminary treatments had little effect upon either sward conditions or animal behaviour.Herbage mass was reduced from 4020 kg dry matter (DM) ha−1to 3290 kg DM ha−1on average over a 5‐d grazing period as a consequence of the relatively low grazing pressure imposed. This resulted in a mean decline in intake per bite of 28%, and the changes for cattle and sheep did not differ significantly. However, there was a marked difference in the other behavioural responses of the two species; in the sheep biting rate fell and grazing time increased with declining herbage mass, particularly in the sward previously grazed infrequently, whereas the changes in the cattle were small.Differences in the botanical composition of the herbage eaten by cattle and sheep were minor, but there was a small but consistent advantage to the sheep in the digestibility of the herbage eaten.In the first of the two periods the variation in surface height after grazing was substantially greater for sheep‐grazed than for cattle‐grazed swards, indicating more patchy grazing by the sheep.Estimates of daily herbage organic matter (OM) intake calculated from ingestive behaviour variables were high (means 38 and 32 g (kg LW)−1for cattle and sheep respectively) and usually declined substantially over a