SUMMARYThis paper deals with theVanguerieae, Cinchoneae, CondamineaeandRondeletieae, and concludes a study on the anatomy of the secundary xylem of theCinchonoideae+Ixoroideae. The taxonomically homogeneousVanguerieaeshow only little variation. Taxonomically theCinchoneae, Condamineae, andRondeletieae, placed near each other in theCinchonoideae, are rather heterogeneous. Their woods show also more anatomical differences, with a similar scala of variation within each tribe. Numerical analysis of the pattern of variation was undertaken. Cluster analysis showed a very distinct bipartition, which remained almost invaried while calculations were executed with different character weighings. This bipartition is not expressed in the existing taxonomic classifications. An attempt to distinguish the three tribes on the basis of their wood anatomy was made, but was unsuccessful.