Pollen of representatives of all genera previously assigned to the ‘Cotuleae’, as well asMinuriafrom the Astereae, is described from studies under light microscopy and scanning and transmission electron microscopy.Cotulas. str.,Leptinella, NanantheaandSolivaare characterized by exines with anthemoid structure; this supports the conclusion that these are natural members of the tribe Anthemideae. The affinities of the non-anthemoid genera are discussed in the light of data on exine structure.Abrotanella, IschneaandCentipedahave senecioid exines, whileCeratogyne, Dimorphocoma, Elachanthus, Isoetopsis, MinuriaandPlagiocheilushave helianthoid exines. Marked differences are shown between the exines ofAbroianellaandIschneawhich do not support the existence of a very close affinity between them. Evidence is provided to support the link betweenIsoetopsisandElachanthusthat has been proposed on the basis of cypsela characters.