The new conodontWebbygnathus munusculumgen. et sp. nov. is described from Eastonian (early Late Ordovician) strata from the central part of the Parkes Zone of the Lachlan Fold Belt and the New England Fold Belt in New South Wales. In the type area south of Gunningbland, central New South Wales, the genus occurs associated with macrofossils of the coral/stromatoporoid assemblage Fauna II (early Eastonian or Ea2); in the New England region it has been obtained from strata on both sides of a major structural feature, the Peel Fault, the associated conodont assemblages indicating an age equivalent to that of coral/stromatoporoid assemblage Fauna III or late Eastonian (Ea3). The apparatus of this species, as presently known, comprises two pectiniform elements, one stellate with a four-rayed basal cavity, the other fundamentally pastinate, with a three-rayed basal cavity.